Additional Functions


Belong­ing to

Gaeabionta  ⇒ Domäne: Eukaryota  ⇒ Reich: Animalia  ⇒ Mittelreich: Eumetazoa  ⇒ Klade: Triploblastica  ⇒ Unterreich: Bilateria  ⇒ Klade: Eubilateria  ⇒ Abzweig: Protostomia  ⇒ Infrareich: Ecdysozoa  ⇒ Superstamm: Panarthropoda  ⇒ Stamm: Arthropoda  ⇒ Unterstamm: Mandibulata  ⇒ Klade: Pancrustacea  ⇒ Unterstamm: Hexapoda  ⇒ Klade: Cercofilata  ⇒ Klasse: Insecta  ⇒ Klade: Dicondylia  ⇒ Klade: Paranotalia  ⇒ Unterklasse: Pterygota  ⇒ Infraklasse: Neoptera  ⇒ ohne Rang: Endoneoptera  ⇒ Überordnung: Coleopterida  ⇒ Ordnung: Coleoptera  ⇒ Unterordnung: Polyphaga  ⇒ Infraordnung: Cucujiformia  ⇒ Überfamilie: Tenebrionoidea  ⇒ Familie: Tenebrionidae

Taxo­nomic seg­ment


This mentioned taxonomy is an attempt to make an conclusive assignment from the different classifications of various scientists. Because the taxonomy may change due to the latest investigative methods and other findings, our map is a guide only.

Name from

Corre­spond­ing author (Name, Year)

Latreille, 1802

Other languages



Lebenszeitraum in Million Jahren (Ma)

Start age: 122.46 Ma - End age: 0.0117 Ma
Upper Pleistocene
Upper/Late Cretaceous
Lower/Early Cretaceous

Additional data by PBDB, License: CC BY

Motility: actively mobile
Environment: terrestrial
Compositon of the remains:


Reference- and Source indication, Literature

Automatic Links to external internet sources (Responsibility is with the operator)

Search for taxonomy at
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Next lower taxonomy level

On the same taxonomic level (siblings) (Count: 25)

Taxonomic assignment (105)

Eukaryota (Domäne)

Animalia (Reich)

Eumetazoa (Mittelreich)


Bilateria (Unterreich)

Eubilateria (Klade)

Protostomia (Abzweig)

Ecdysozoa (Infrareich)

Panarthropoda (Superstamm)

Arthropoda (Stamm)

Mandibulata (Unterstamm)

Pancrustacea (Klade)

Hexapoda (Unterstamm)

Cercofilata (Klade)

Insecta (Klasse)

Dicondylia (Klade)

Paranotalia (Klade)

Pterygota (Unterklasse)

Neoptera (Infraklasse)

Endoneoptera (ohne Rang)

Coleopterida (Überordnung)

Coleoptera (Ordnung)

Polyphaga (Unterordnung)

Cucujiformia (Infraordnung)

Tenebrionoidea (Überfamilie)

Tenebrionidae (Familie)

Tenebrioninae (Unterfamilie)

Alphitobiini (Tribus)

Alphitopsis (Gattung)


Amarygmini (Tribus)

Cymatothes (Gattung)


Meracantha (Gattung)


Amphidorini (Tribus)

Eleodes (Gattung)

Eleodes (Blapylis) (Untergattung)


Eleodes (Cratidus) (Untergattung)


Eleodes (Eleodes) (Untergattung)




Eleodes (Litheleodes) (Untergattung)


Eleodes (Melaneleodes) (Untergattung)



Eleodes (Xysta) (Untergattung)


Blaptini (Tribus)

Blaptina (Untertribus)

Blaps (Gattung)

Bolitophagini (Tribus)

Bolitophagus (Gattung)


Eledona (Gattung)


Proteleates (Gattung)


Rhipidandrus (Gattung)


Eulabini (Tribus)

Apsena (Gattung)




Eulabis (Gattung)

Helopini (Tribus)

Cylindrinotina (Untertribus)

Nalassus (Gattung)

Helopina (Untertribus)

Cryptohelops (Gattung)


Helops (Gattung)



Stenohelops (Gattung)



Melanimonini (Tribus)

Melanimon (Gattung)


Opatrini (Tribus)

Blapstinina (Untertribus)

Blapstinus (Gattung)


Ulus (Gattung)

Neopachypterina (Untertribus)

Eupachypterus (Gattung)


Opatrina (Untertribus)

Ephalus (Gattung)


Gonocephalum (Gattung)


Opatrum (Gattung)

Palaeosclerum (Gattung)


Palorini (Tribus)

Palorus (Gattung)


Vabole (Gattung)


Pedinini (Tribus)

Leichenina (Untertribus)

Leichenum (Gattung)

Scotobiini (Tribus)

Scotobius (Gattung)


Tenebrionini (Tribus)

Tenebrio (Gattung)





Titaenini (Tribus)

Pseudhelops (Gattung)


Toxicini (Tribus)

Dysantina (Untertribus)

Wattius (Gattung)


Triboliini (Tribus)

Hypogena (Gattung)


Tribolium (Gattung)

Ulomini (Tribus)

Uloma (Gattung)


Refresh list

Actuality: 23. Sep 2023 - 17:55:27

References by PBDB, License: CC BY

  • M. E. Clapham, 2020 - Insect taxonomy information from the internet, with fixing of species orphaned by genus synonymy - (), - (unpublished, English)
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  • M. V. Nabozhenko, 2019 - The fossil record of darkling beetles (Insecta: Coleoptera: Tenebrionidae) - Geosciences (9), 1-20 (journal article, English)
  • Y. Bousquet, 2018 - Catalogue of Tenebrionidae (Coleoptera) of North America - ZooKeys (728), 1-455 (serial monograph, English)
  • M. Nabozhenko, 2017 - The oldest opatrine terrestrial darkling beetle (Coleoptera: Tenebrionidae: Tenebrioninae) from the Paleocene of Menat (France) - PalZ (91), 307-313 (journal article, English)
  • V. I. Alekseev, 2017 - Palorus platycotyloides sp. n., the second fossil representative of the tribe Palorini (Coleoptera: Tenebrionidae) from Baltic amber - Acta Zoologica Bulgarica (69), 167-170 (journal article, English)
  • L. C. Ramírez, 2016 - First fossil assemblage of darkling beetles (Coleoptera: Tenebrionidae) from Buenos Aires, Argentina: evidence of dry climate during the Late Pleistocene in the humid Pampas - Annales Zoologici (66), 643-652 (journal article, English)
  • M. V. Nabozhenko, 2016 - A new species of the genus Nalassus Mulsant (Coleoptera: Tenebrionidae: Helopini) from the Baltic amber - Paleontological Journal (50), 947-952 (journal article, English)
  • V. I. Alekseev, 2015 - A new fossil tenebrionid beetle of the tribe Palorini (Coleoptera: Tenebrionidae) from Eocene Baltic amber - The Coleopterists Society Monograph (14), 127-130 (journal article, English)
  • M. V. Nabozhenko, 2014 - Cryptohelops menaticus – a new genus and species of the tribe Helopini (Coleoptera: Tenebrionidae) from the Palaeocene of Menat (France) - Comptes Rendus Palevol (13), 65-71 (journal article, English)
  • A. G. Kirejtshuk, 2011 - First Mesozoic representative of the subfamily Tenebrioninae (Coleoptera, Tenebrionidae) from the Lower Cretaceous of Yixian (China, Liaoning) - Entomological Review (92), 97-100 (journal article, English)
  • P. Bouchard, 2011 - Family-group names in Coleoptera (Insecta) - ZooKeys (88), 1-972 (journal article, English)
  • A. G. Kirejtshuk, 2010 - New genus and species of the tribe Opatrini (Coleoptera, Tenebrionidae, Tenebrioninae) from the lowermost Eocene amber of Paris Basin - Proceedings of the Zoological Institute RAS (314), 191-196 (journal article, English)
  • A. G. Kirejtshuk, 2008 - A new species of the genus Pentaphyllus Dejean, 1821 (Coleoptera, Tenebrionidae, Diaperinae) from the Baltic amber and checklist of the fossil Tenebrionidae - Zoosystematica Rossica (17), 131-137 (journal article, English)
  • P. Bouchard, 2007 - Nomenclatural notes on tenebrionid beetles of the Palaearctic region (Insecta: Coleoptera) - Annales Zoologici (57), 385-394 (journal article, English)
  • C. P. Green, 2006 - Marine Isotope Stage 9 environments of fluvial deposits at Hackney, north London, UK - Quaternary Science Reviews (25), 89-113 (journal article, English)
  • P. Bouchard, 2005 - Synoptic classification of the world Tenebrionidae (Insecta: Coleoptera) with a review of family-group names - Annales Zoologici (55), 499-530 (journal article, English)
  • E. G. Matthews, 2003 - Ulomotypus Broun a member of the new subfamily Palorinae, with remarks on Aphtora Bates and Demtrius Broun (Coleoptera, Tenebrionidae) - New Zealand Entomologist (26), 7-14 (journal article, English)
  • J. T. Doyen, 1994 - Tenebrionidae from Dominican amber (Coleoptera) - Entomologica Scandinavica (25), 27-51 (journal article, English)
  • J. C. Watt, 1992 - Tenebrionidae (Insecta: Coleoptera): catalogue of types and keys to taxa - Fauna of New Zealand (26), 1-70 (serial monograph, English)
  • W. D. Pierce, 1954 - Fossil arthropods of California no. 19. The Tenebrionidae-Scaurinae of the asphalt deposits - Bulletin of the Southern California Academy of Sciences (53), 93-98 (journal article, English)
  • H. Haupt, 1950 - Die Käfer (Coleoptera) aus der eozänen Braunkohle des Geiseltales - Geologica (6), 1-168 (journal article, German)
  • E. Reitter, 1922 - Bestimmungstabelle der palaearktischen Helopinae (Col. Tenebrionidae.) I. Teil - Wiener Entomologische Zeitung (39), 1-44 (journal article, German)
  • H. F. Wickham, 1914 - New Miocene Coleoptera from Florissant - Bulletin of the Museum of Comparative Zoology, Harvard College (53), 423-494 (serial monograph, English)
  • H. F. Wickham, 1914 - Twenty new Coleoptera from the Florissant Shales - Transactions of the American Entomological Society (40), 257-270 (journal article, English)
  • H. F. Wickham, 1913 - Fossil Coleoptera from Florissant in the United States National Museum - Proceedings of the United States National Museum (45), 283-303 (journal article, English)
  • F. E. Blaisdell, 1909 - A monographic revision of the Coleoptera belonging to the tenebrionide tribe Eleodiini inhabiting the United States, Lower California, and adjacent islands - Bulletin of the United States National Museum (63), 1-524 (serial monograph, English)
  • H. F. Wickham, 1909 - New Fossil Coleoptera from Florissant - American Journal of Science, Series 4 (28), 126-130 (journal article, English)
  • F. Grinnell, 1908 - Quaternary myriopods and insects of California - University of California Publications, Bulletin of the Department of Geology (5), 207-215 (journal article, English)
  • A. Fauvel, 1905 - Faune analytique des coléoptères de la Nouvelle-Calédonie. 3e partie (1) Tenebrionidae (Suite) - Revue d’Entomologie (24), 209-244 (book, French)
  • S. H. Scudder, 1900 - Adephagous and clavicorn Coleoptera from the Tertiary deposits at Florissant, Colorado with descriptions of a few other forms and a systematic list of the non-rhynchophorus Tertiary Coleoptera of North America - Monographs of the United States Geological Survey (40), 1-148 (serial monograph, English)
  • T. L. Casey, 1891 - Coleopterological Notices. III - Annals of the New York Academy of Sciences (6), 9-214 (journal article, English)
  • S. H. Scudder, 1879 - Appendix A. The fossil insects collected in 1877, by Mr. G.M. Dawson, in the interior of British Columbia - Geological Survey of Canada, Report of Progress for 1877-1878 (), 175-185 (book chapter, English)
  • C. von Heyden, 1866 - Käfer und Polypen aus der Braunkohle des Siebengebirges - Palaeontographica (15), 131-156 (journal article, German)
  • C. von Heyden, 1862 - Gliederthiere aus der Braunkohle des Niederrhein's, der Wetterau und der Röhn - Palaeontographica (10), 62-82 (journal article, German)
  • C. Von Heyden, 1859 - Fossile Insekten aus der Rheinischen Braunkhohle - Palaeontographica (8), 1-15 (journal article, German)
  • E. Mulsant, 1853 - Essai d’une division des derniers Mélasomes - Mémoires de l’Académie Nationale des Sciences, Belles-Lettres et Arts de Lyon. Classe des Sciences. Nouvelle Série (2), 226-329 (journal article, French)
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GUSID (Global unique identifier short form) R63x9PBrKkeRGzE-ewbv9A
GUID (Global unique identifier) F4F1AD47-6BF0-472A-911B-313E7B06EFF4
Database ID 22155