† Capitosauria


Additional Functions


Belong­ing to

Gaeabionta  ⇒ Domäne: Eukaryota  ⇒ Reich: Animalia  ⇒ Mittelreich: Eumetazoa  ⇒ Klade: Triploblastica  ⇒ Klade: Nephrozoa  ⇒ Abzweig: Deuterostomia  ⇒ Stamm: Chordata  ⇒ Unterstamm: Vertebrata  ⇒ Infrastamm: Gnathostomata  ⇒ Superklasse: Tetrapoda  ⇒ Klasse: Amphibia  ⇒ Ordnung: Temnospondyli  ⇒ ohne Rang: Eutemnospondyli  ⇒ ohne Rang: Eryopiformes  ⇒ ohne Rang: Stereospondylomorpha  ⇒ ohne Rang: Stereospondyli

Taxo­nomic seg­ment

ohne Rang

This mentioned taxonomy is an attempt to make an conclusive assignment from the different classifications of various scientists. Because the taxonomy may change due to the latest investigative methods and other findings, our map is a guide only.

Name from

Corre­spond­ing author (Name, Year)

Yates & Warren, 2000

Other languages



Lebenszeitraum in Million Jahren (Ma)

Start age: 251.2 Ma - End age: 201.3 Ma
Lower/Early Jurassic
Upper/Late Triassic
(Obere Trias)
Middle Triassic
(Mittlere Trias)
Lower/Early Triassic
(Untere Trias)

Additional data by PBDB, License: CC BY

Motility: actively mobile
Habit: amphibious
Environment: brackish, freshwater, terrestrial
Compositon of the remains:


Reference- and Source indication, Literature

Automatic Links to external internet sources (Responsibility is with the operator)

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Next lower taxonomy level

Gattung: Benthosuchus Efremov, 1936

Gattung: Calmasuchus Fortuny et al., 2011

Überfamilie: Capitosauroidea Watson, 1919

Gattung: Edingerella Schoch & Milner, 2000

Familie: Sclerothoracidae Huene, 1932

Gattung: Tomeia Eltink et al., 2017

Gattung: Vladlenosaurus Morkovin & Novikov, 2000

Gattung: Warrenisuchus Maganuco et al., 2009

Gattung: Watsonisuchus Ochev, 1966

Gattung: Wetlugasaurus Riabinin, 1930

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Actuality: 20. Feb 2024 - 19:54:17

On the same taxonomic level (siblings) (Count: 12)

Taxonomic assignment (139)

Eukaryota (Domäne)

Animalia (Reich)

Eumetazoa (Mittelreich)


Nephrozoa (Klade)

Deuterostomia (Abzweig)

Chordata (Stamm)

Vertebrata (Unterstamm)

Gnathostomata (Infrastamm)

Tetrapoda (Superklasse)

Amphibia (Klasse)

Temnospondyli (Ordnung)

Eutemnospondyli (ohne Rang)

Eryopiformes (ohne Rang)


Stereospondyli (ohne Rang)

Capitosauria (ohne Rang)

Benthosuchus (Gattung)






Calmasuchus (Gattung)


Capitosauroidea (Überfamilie)

Abiadisaurus (Gattung)


Antarctosuchus (Gattung)


Cyclotosauridae (Familie)

Cyclotosaurinae (Unterfamilie)

Heylerosaurinae (Unterfamilie)



Stenotosaurinae (Unterfamilie)

Cyclotosaurus (Gattung)









Deltacephalidae (Familie)

Eocyclotosaurus (Gattung)






Eryosuchidae (Familie)

Eryosuchus (Gattung)



Heylerosauridae (Familie)

Rhadalognathus (Gattung)


Yuanansuchus (Gattung)




Bulgosuchus (Gattung)


Cherninia (Gattung)



Heptasaurus (Gattung)

Jammerbergia (Gattung)


Kestrosaurus (Gattung)



Komatosuchus (Gattung)


Parotosuchus (Gattung)










Selenocarinae (Unterfamilie)

Samarabatrachus (Gattung)


Selenocara (Gattung)



Stenotosaurus (Gattung)






Tatrasuchus (Gattung)



Trochanterium (Gattung)


Wellesaurus (Gattung)


Xenotosuchus (Gattung)


Mastodonsaurus (Gattung)





Mentosaurus (Gattung)


Meyerosuchus (Gattung)


Odenwaldia (Gattung)











Parotosuchidae (Familie)



Rewanobatrachus (Gattung)

Yarengia (Gattung)


Yarengiidae (Familie)

Edingerella (Gattung)



Sclerothorax (Gattung)


Tomeia (Gattung)


Vladlenosaurus (Gattung)


Warrenisuchus (Gattung)


Watsonisuchus (Gattung)




Wetlugasaurus (Gattung)







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Actuality: 20. Feb 2024 - 19:54:16

Occurrences (2)

Refresh list | Fold level: 1 - 2 - 3 - 4 - 5 - Unfold allActuality: 20. Feb 2024 - 19:54:16

Locations with GPS information

References by PBDB, License: CC BY

  • R. R. Schoch, 2018 - The temnospondyl Parotosuchus nasutus (v. Meyer, 1858) from the Early Triassic Middle Buntsandstein of Germany - Palaeodiversity (11), 107-126 (journal article, English)
  • B. R. Peecook, 2018 - Updated geology and vertebrate paleontology of the Triassic Ntawere Formation of northeastern Zambia, with special emphasis on the archosauromorphs - Society of Vertebrate Paleontology Memoir 17 (37), 8-38 (journal article, English)
  • E. Eltink, 2017 - A capitosauroid from the Lower Triassic of South America (Sanga do Cabral Supersequence: Paraná Basin), its phylogenetic relationships and biostratigraphic implications - Historical Biology (29), 863-874 (journal article, English)
  • M. Marzola, 2017 - Cyclotosaurus naraserluki, sp. nov., a new Late Triassic cyclotosaurid (Amphibia, Temnospondyli) from the Fleming Fjord Formation of the Jameson Land Basin (East Greenland) - Journal of Vertebrate Paleontology (37), e1303- (journal article, English)
  • J. Liu, 2016 - Yuanansuchus maopingchangensis sp. nov., the second capitosauroid temnospondyl from the Middle Triassic Badong Formation of Yuanan, Hubei, China - PeerJ (4), e1903- (journal article, English)
  • A. Dahoumane, 2016 - A new Mastodonsauroid Temnospondyl from the Triassic of Algeria: Implications for the biostratigraphy and palaeoenvironments of the Zarzaïtine Series, northern Sahara - Comptes Rendus Palevol (15), 918-926 (journal article, English)
  • I. V. Novikov, 2016 - New tempospondyl amphibians from the basal Triassic of the Obshchii Syrt Highland, Eastern Europe - Paleontological Journal (50), 297-310 (journal article, English)
  • F. Witzmann, 2016 - A new species of Cyclotosaurus (Stereospondyli, Capitosauria) from the Late Triassic of Bielefeld, NW Germany, and the intrarelationships of the genus - Fossil Record (19), 83-100 (journal article, English)
  • L. F. Rinehart, 2015 - Eocyclotosaurus appetolatus, a new cyclotosaurid amphibian from the Middle Triassic (Perovkan) Moenkopi Formation of New Mexico, U.S.A. - Journal of Vertebrate Paleontology (35), e9291- (journal article, English)
  • C. A. Sidor, 2014 - A new capitosauroid temnospondyl from the Middle Triassic upper Fremouw Formation of Antarctica - (34), 539-548 (journal article, English)
  • R. R. Schoch, 2013 - The evolution of major temnospondyl clades: an inclusive phylogenetic analysis - Journal of Systematic Palaeontology (), - (journal article, English)
  • I. V. Novikov, 2012 - New data on trematosauroid labyrinthodonts of Eastern Europe: 4. Genus Benthosuchus Efremov, 1937 - Paleontological Journal (46), 68-79 (journal article, English)
  • J. Fortuny, 2011 - A new capitosaur from the Middle Triassic of Spain and the relationships within the Capitosauria - Acta Palaeontologica Polonica (56), 553-566 (journal article, English)
  • R. R. Schoch, 2011 - How diverse is the temnospondyl fauna in the Lower Triassic of southern Germany? - Neues Jahrbuch für Geologie und Paläontologie Abhandlungen (261), 49-60 (journal article, English)
  • T. Sulej, 2011 - A new large capitosaur temnospondyl amphibian from the Early Triassic of Poland - Acta Palaeontologica Polonica (58), 65-75 (journal article, English)
  • S. Maganuco, 2009 - An exquisite specimen of Edingerella madagascariensis (Temnospondyli) from the Lower Triassic of NW Madagascar; cranial anatomy, phylogeny, and restorations - Memorie della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano (36), 1-72 (journal article, English)
  • M. A. Shishkin, 2009 - The Early Triassic temnospondyls of the Czatkowice 1 tetrapod assemblage - Palaeontologia Polonica (65), 31-77 (journal article, English)
  • M. Moser, 2007 - Revision of the type material and nomenclature of Mastodonsaurus giganteus (Jaeger) (Temnospondyli) from the Middle Triassic of Germany - Palaeontology (50), 1245-1266 (journal article, English)
  • R. R. Schoch, 2007 - Anatomy and relationships of the Triassic temnospondyl Sclerothorax - Acta Palaeontologica Polonica (52), 117-136 (journal article, English)
  • T. Sulej, 2005 - The temnospondyl amphibian Cyclotosaurus from the Upper Triassic of Poland - Palaeontology (48), 157-170 (journal article, English)
  • J. Liu, 2005 - The first complete mastodonsauroid skull from the Triassic of China: Yuanansuchus laticeps gen. et sp. nov. - Journal of Vertebrate Paleontology (25), 725-728 (journal article, English)
  • S. J. Nesbitt, 2005 - The Moenkopi Formation along the Little Colorado River in eastern Arizona - Guidebook to the Triassic Formations of the Colorado Plateau in northern Arizona, Mesa Southwest Museum Bulletin (9), 13-23 (book/book chapter, English)
  • M. A. Shishkin, 2004 - Re-evaluation of Kestrosaurus Haughton, a capitosaurid temnospondyl amphibian from the Upper Beaufort Group of South Africa - Russian Journal of Herpetology (11), 121-138 (journal article, English)
  • V. P. Tverdokhlebov, 2003 - Tetrapod localities from the Triassic of the SE of European Russia - Earth-Science Reviews (60), 1-66 (journal article, English)
  • R. J. Damiani, 2003 - A review of the South African temnospondyl amphibian record - Palaeontologia africana (39), 21-36 (journal article, English)
  • R. J. Damiani, 2003 - New mastodonsaurid temnospondyls from the Cynognathus Assemblage Zone (Upper Beaufort Group; Karoo Basin) of South Africa - Journal of Vertebrate Paleontology (23), 54-66 (journal article, English)
  • J. S. Steyer, 2003 - A revision of the early Triassic “capitosaurs” (Stegocephali, Stereospondyli) from Madagascar, with remarks on their comparative ontogeny - Journal of Vertebrate Paleontology (23), 544-555 (journal article, English)
  • M. Morales, 2002 - A re-assessment of Parotosuchus africanus (Broom), a capitosauroid temnospondyl amphibian from the Triassic of South Africa - Journal of Vertebrate Paleontology (22), 1-11 (journal article, English)
  • R. J. Damiani, 2001 - A systematic revision and phylogenetic analysis of Triassic mastodonsauroids (Temnospondyli: Stereospondyli) - Zoological Journal of the Linnean Society (133), 379-482 (journal article, English)
  • R. Schoch, 2000 - Stereospondyli - Handbuch der Paläoherpetologie - Encyclopedia of Paleoherpetology (3B), 1-203 (book/book chapter, English)
  • M. A. Shishkin, 2000 - Permian and Triassic temnospondyls from Russia - The Age of Dinosaurs in Russia and Mongolia. Cambridge University Press, Cambridge (), 35-59 (book/book chapter, English)
  • R. R. Schoch, 2000 - The status and osteology of two new cyclotosaurid amphibians from the Upper Moenkopi Formation of Arizona (Amphibia: Temnospondyli; Middle Triassic) - Neues Jahrbuch für Geologie und Paläontologie Abhandlungen (216), 387-411 (journal article, English)
  • A. M. Yates, 2000 - The phylogeny of the ‘higher’ temnospondyls (Vertebrata: Choanata) and its implications for the monophyly and origins of the Stereospondyli - Zoological Journal of the Linnean Society (128), 77-121 (journal article, English)
  • S. G. Lucas, 2000 - Biochronological significance of Triassic nonmarine tetrapod records from marine strata - Albertiana (24), 27-32 (journal article, English)
  • R. R. Schoch, 1999 - The Triassic labyrinthodonts from Germany - Zentralblatt für Geologie und Paläontologie Teil I (7-8), 629-650 (journal article, English)
  • R. J. Damiani, 1999 - Giant temnospondyl amphibians from the Early to Middle Triassic Narrabeen Group of the Sydney Basin, New South Wales, Australia - Alcheringa (23), 87-109 (journal article, English)
  • R. R. Schoch, 1999 - Comparative osteology of Mastodonsaurus giganteus (Jaeger, 1828) from the Middle Triassic (Lettenkeuper: Longobardian) of Germany (Baden-Württemberg, Bayern, Thüringen) - Stuttgarter Beiträge zur Naturkunde Serie B (278), 1-170 (journal article, English)
  • R. N. Mukherjee, 1998 - New capitosaurid amphibians from the Triassic Denwa Formation of the Satpura Gondwana Basin, central India - Ameghiniana (22), 317-327 (journal article, English)
  • R. R. Schoch, 1997 - A new capitosaur amphibian from the Upper Lettenkeuper (Triassic: Ladinian) of Kupferzell (Southern Germany) - Neues Jahrbuch für Geologie und Paläontologie (203), 239-272 (journal article, English)
  • M. A. Shishkin, 1996 - A new lydekkerinid (Amphibia, Temnospondyli) from the lower Triassic of South Africa: implications for evolution of the early capitosauroid cranial pattern - Philosophical Transactions of the Royal Society of London B (351), 1635-1659 (journal article, English)
  • I. V. Novikov, 1992 - New Middle Triassic labyrinthodonts from the Pechora Urals - Paleontological Journal (26), 92-102 (journal article, English)
  • R. L. Carroll, 1988 - - Vertebrate Paleontology and Evolution (), 1-698 (compendium, English)
  • A. A. Warren, 1988 - A new capitosaurid amphibian from the early Triassic of Queensland, and the ontogeny of the capitosaur skull - Palaeontology (31), 857-876 (journal article, English)
  • M. Morales, 1984 - Odenwaldia heidelbergensis, a new benthosuchid stegocephalian from the Middle Buntsandstein of the Odenwald, Germany - Neues Jahrbuch für Geologie und Paläontologie Monatshefte (1984), 673-683 (journal article, English)
  • D. Kamphausen, 1983 - Stenotosaurus gracilis, ein neuer Capitosauride (Stegocephalia) aus den Unteren Röttonen Oberfrankens - Neues Jahrbuch für Geologie und Paläontologie, Monatshefte (1983), 119-128 (journal article, German)
  • R. Ingavat, 1981 - Cyclotosaurus cf. posthumus Fraas (Capitosauridae, Stereospondyli) from the Huai Hin Lat Formation (Upper Triassic), Northeastern Thailand, with a note on capitosaurid biogeography - Géobios (14), 711-725 (journal article, English)
  • A. A. Warren, 1980 - Parotosuchus from the Early Triassic of Queensland and Western Australia - Alcheringa (4), 25-36 (journal article, English)
  • C. Yang, 1978 - [A Late Triassic vertebrate fauna from Fukang, Sinkiang] - Memoirs of the Institute of Vertebrate Paleontology and Paleoanthropology, Academia Sinica (13), 60-67 (journal article, Chinese)
  • S. Chernin, 1978 - Three capitosaurs from the Triassic of South Africa: Parotosuchus africanus (Broom 1909); Kestrosaurus dreyeri Haughton 1925, and Parotosuchus dirus sp. nov. - Palaeontologia africana (21), 79-100 (journal article, English)
  • R. L. Paton, 1976 - A replacement name for Parotosaurus Jaekel (Amphibia; Labyrhinthodontia) - Palaeontology (19), 415-416 (journal article, English)
  • S. Chernin, 1975 - Further considerations of the capitosaurids from the Upper Luangwa Valley, Zambia - Palaeontologia africana (18), 143-148 (journal article, English)
  • J. W. Cosgriff, 1972 - Parotosaurus wadei, a new capitosaurid from New South Wales - Journal of Paleontology (46), 545-555 (journal article, English)
  • J.-P. Lehman, 1971 - Nouveaux vertebres fossiles du Trias de la Serie de Zarzaitine - Annales de Paleontologie (57), 71-116 (journal article, French)
  • O. Kuhn, 1971 - - Die Saurier der deutschen Trias [Reptiles of the German Triassic] (), 1-105 (book, German)
  • A. A. Howie, 1970 - A new capitosaurid labyrinthodont from East Africa - Palaeontology (13), 210-253 (journal article, English)
  • T. R. Chowdhury, 1970 - A new capitosaurid amphibian from the Triassic Yerrapalli Formation fo the Panhita-Godavari Valley - Journal of the Geological Society of India (11), 155-162 (journal article, English)
  • C.-C. Young, 1966 - On the first discovery of capitosaurid from Sinkiang - Vertebrata PalAsiatica (10), 58-62 (journal article, Chinese)
  • S. P. Welles, 1965 - A revision of the labyrinthodont family Capitosauridae, and a description of Parotosaurus peabody, n. sp. from the Wupatki Member of the Moenkopi Formation of northern Arizona - University of California Publications in Geological Sciences (54), 1-148 (serial monograph, English)
  • D. M. S. Watson, 1962 - The evolution of the labyrinthodonts - Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences (245), 219-265 (journal article, English)
  • J.-P. Lehman, 1961 - Les stégocéphales du Trias de Madagascar - Annales de Paléontologie (47), 109-154 (journal article, French)
  • M. A. Shishkin, 1960 - [Yarengiidae, a new family of Triassic labyrinthodonts] - Paleontological Journal (1), 97-106 (journal article, English)
  • D. M. S. Watson, 1958 - A new labyrinthodont (Paracyclotosaurus) from the upper Trias of New South Wales - Bulletin of the British Museum (Natural History) Geology (3), 233-262 (journal article, English)
  • A. S. Romer, 1947 - Review of the Labyrinthodontia - Bulletin of the Museum of Comparative Zoology (99), 1-368 (journal article, English)
  • S. P. Welles, 1947 - Vertebrates from the Upper Moenkopi Formation of northern Arizona - University of California Publications in Geological Sciences (27), 241-294 (journal article, English)
  • J. A. Efremov, 1940 - Predvaritel'noye opisaniye novykh form permskoy i triasovoy fauny nazemnykh pozvonochnykh SSSR [Preliminary description of new forms from the Permian and Triassic terrestrial vertebrate fauna of the USSR] - Trudy Paleontologicheskogo Instituta Akademiy Nauk SSSR (10), 1-140 (journal article, Russian)
  • F. Broili, 1937 - Beobachtungen an Wirbeltieren der Karrooformation. XXVII. Über einen Capitosauriden aus der Cynognathus-Zone - Sitzungsberichte der Bayerischen Akademie der Wissenschaften, Mathematisch-naturwissenschaftliche Abteilung (1937), 97-117 (journal article, German)
  • G. Säve-Söderbergh, 1935 - On the dermal bones of the head in labyrinthodont stegocephalians and primitive Reptilia, with special reference to Eotriassic stegocephalians from East Greenland - Meddelelser om Grønland (98), 1-211 (serial monograph, English)
  • B. Brown, 1933 - A new genus of Stegocephalia from the Triassic of Arizona - American Museum Novitates (640), 1-4 (journal article, English)
  • F. v. Huene, 1932 - Ein neuartiger Stegocephalen-Fund aus dem oberhessischen Buntsandstein [A new stegocephalian specimen from the Upper Hesse Buntsandstein] - Palaeontologische Zeitschrift (14), 200-229 (journal article, German)
  • O. Kuhn, 1932 - Labyrinthodonten und Parasuchier aus dem mittleren Keuper von Ebrach in Oberfranken - Neues Jahrbuch für Mineralogie, Geologie und Paläontologie (69), 94-144 (journal article, English)
  • S. H. Haughton, 1925 - Investigations in South African fossil reptiles and amphibians (Part 13) - Annals of the South African Museum (22), 227-261 (journal article, English)
  • D. M. S. Watson, 1919 - The structure, evolution and origin of the Amphibia.—The "orders" Rachitomi and Stereospondyli - Philosophical Transactions of the Royal Society of London, Series B (209), 1-73 (journal article, English)
  • W. J. Sinclair, 1917 - A new labyrinthodont from the Triassic of Pennsylvania - American Journal of Science (256), 319-321 (journal article, English)
  • C. Wiman, 1914 - Über die Stegocephalen aus der Trias Spitzbergens - Bulletin of the Geological Institutions of the University of Uppsala (13), 1-34 (journal article, German)
  • O. Jaekel, 1914 - Über die Wirbeltierfunde in der oberen Trias von Halberstadt [On the vertebrate remains in the Upper Triassic of Halberstadt] - Paläontologische Zeitschrift (1), 155-215 (journal article, German)
  • H. Schroeder, 1913 - Ein Stegocephalen-Schädel von Helgoland - Jahrbuch der Preussischen Geologischen Landesanstalt (33), 232-264 (journal article, German)
  • R. Broom, 1909 - Notice of some New South African Fossil Amphibians and Reptiles - Annals of the South African Museum (7), 270-278 (journal article, English)
  • A. S. Woodward, 1904 - On two new labyrinthodont skulls of the genera Capitosaurus and Aphaneramma - Proceedings of the Zoological Society of London (1904), 170-176 (journal article, English)
  • F. v. Huene, 1902 - Übersicht über die Reptilien der Trias [Review of the Reptilia of the Triassic] - Geologische und Paläontologische Abhandlungen (Neue Serie). Gustav Fischer Verlag, Jena (6), 1-84 (serial monograph, German)
  • H. G. Seeley, 1876 - On the posterior portion of a lower jaw of Labyrinthodon (L. Lavisi), from the Trias of Sidmouth - Quarterly Journal of the Geological Society of London (32), 278-284 (journal article, English)
  • R. Owen, 1861 - - Palaeontology, or a Systematic Summary of Extinct Animals and their Geological Relations. Second Edition. Adam and Charles Black, Edinburgh (), 1-463 (book, English)
  • H. v. Meyer, 1855 - - Zur Fauna der Vorwelt. Die Saurier des Muschelkalkes mit Rücksicht auf die Saurier aus Buntem Sandstein und Keuper [The Fauna of the Ancient World. The Saurians of the Muschelkalk with Regard to the Saurians of the Buntem Sandstone and Keuper] (), viii-167 (book, German)
  • H. v. Meyer, 1845 - System der fossilen Saurier [Taxonomy of fossil saurians] - Neues Jahrbuch für Mineralogie, Geognosie, Geologie und Petrefakten-Kunde (1845), 278-285 (journal article, German)
  • G. Münster, 1836 - Communications addressed to Professor Bronn - Neues Jahrbuch für Mineralogie, Geognosie, Geologie und Petrefaktenkunde (1836), 580-581 (journal article, German)
  • F. Holl, 1829 - - Handbuch der Petrefactenkunde, mit einer Einleitung über die Vorwelt der organischen Wesen auf der Erde von Dr. Ludwig Choulant (1), 1-115 (serial monograph, German)
  • G. F. Jaeger, 1828 - Über die Fossile Reptilien, Welche in Württemberg Aufgefunden Sind [On the Fossil Reptiles That Are Found in Württemberg] - Verlag der Metzler'schen Buchhandlung, Stuttgart (), 1-48 (book, German)


GUSID (Global unique identifier short form) GdnqcnN_uEiyPCF0Pz8WuA
GUID (Global unique identifier) 72EAD919-7F73-48B8-B23C-21743F3F16B8
Database ID 5576